Important point here that mail (in this case over SMTP) was basically the earliest internet application that had to "just work" because it was fire-and-forget. FTP came before mail, and was critical, but also it was something that when it failed, it failed noisily and you could call someone to fix it. If your mail is not delivered, you might never know!
Oops, forgot to link to the full interview.
https://www.computerhistory.org/collections/catalog/102738738